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The plenum chamber is a chamber located in front of the windshield, where air pressure is above atmospheric. It provides a volume of air which is drawn into the car from the chamber through the heater, air-conditioner or vents. Early cars had a vent in front of the windshield and this could be opened by a lever in the cabin to allow air into the cabin. Later cars with a plenum chamber had a grille, which was the opening to the chamber.
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