Toyota Land Cruiser vs Land Rover Defender

What's the difference?

The Toyota LandCruiser 300 GR Sport is a total knockout when it comes to its looks, comfort and practicality. It’s stupidly easy to drive and has a fuel efficiency that should make you swoon but the servicing is a bit annoying and it’s not the most affordable option on the market. However, even though it hasn't benefited from any updates this year, you don't feel like you're missing out on any luxuries.

My son adores it, wants it and will be very disappointed when we hand it back. Naturally, he gives it a 10/10.

The Land Rover Defender in its current form is a revelation in terms of, well, everything. The 110 is refined, sure-footed and comfortable on-road and it’s more capable off-road than ever before – and assuredly so.

It’s wholeheartedly embraced positive change – in terms of creature comforts, driveability and safety – and has lost none of the traditional Defender spirit.

The new Defender has managed to satisfy (placate?) die-hard fans and it’s attracted a whole bunch of new ones at the same time.

The 75th anniversary treatment doesn’t add anything of substance to the Defender package, but it doesn’t need to – that’s not the point – and to Landie lovers it’s all cream on top. 

Design is 100 per cent subjective. I love the look of the LC300 GR Sport because it features what I feel is a timeless, handsome shape.

For some, it may look too big and brutish, because you could throw a bunch of beefy words at it, and somehow, they’d all fit the roadside presence of this extra-large SUV.

What visually sets the GR Sport apart from its siblings are the multitude of black accents across the body from the rear badging, wheels and arches, door handles and roof racks. The grille is also distinctly different in its horizontal design with the Toyota badging capitalised in a bold white font for extra effect. 

Head inside and the cabin looks ruggedly capable with the dimensions of the centre console and dashboard complementing the external looks.

Technology looks upmarket, joinery is solidly put together and the leather upholstery adds to the plush comfort of the seats but there is a sense of capability in the physical buttons and dials still found on the dashboard.

Which is where I start to really like the LC300 GR Sport because everything is where you expect it to be - this is not a car you’ll get flustered in but nor does it have that generic Toyota look.

The new Defender manages to balance the distinctive shape and spirit of the old-school Defender with the new-generation’s style and presence – and the striking Grasmere Green exterior of this 75th anniversary variant and its interior touches all complement that blend.

This is unmistakably a Defender but one that’s been dragged kicking and screaming into the 21st century, whacked in the face and torso by modern styling cues, while still retaining the heart of adventure these Landies have always been renowned for.

Just one thing about this special-edition Defender, though. The Grasmere Green is certainly an eye-catching colour, but that’s a ‘for better or worse’ scenario.

Mostly worse – for me anyway – because I noticed that during my time in this Landie I received quite a lot of annoyed eye-rolls from other drivers and passengers in traffic and I reckon the polarising colour was the culprit – or maybe they were just eye-rolling at me?

No matter, because if you can afford one of these I don’t think you’ll be at all concerned about what other people think of you…

The front row is roomy and even with a co-pilot, you feel like you have yards of space to settle into. The seats offer a wide seat and plush padding, as well as heat and ventilation functions which adds comfort for longer trips.

Individual storage is good with a large glove box, two cupholders, two drink bottle holders and a large middle console that also features a 'cool box' function for drinks or snacks.

The comfort of the front is replicated in the backseat, which deftly accepts the height of my 183cm father. The width of the seat can easily accommodate three adults in relative comfort or, if you have a few tots in tow, three child seats.

Amenities and storage in the back row are what you'd expect for a top model in this class. The rear outboard seats feature heat and ventilation functions and there is dual air-conditioning and climate control, as well as, four directional air vents.

The fold-down armrest features two cupholders while the doors hold a drink bottle each and a small storage bin. There are also two map pockets and you can access the middle console cool box with a rear-mounted button.

The GR Sport does remind you that it's a big car with its 235mm ground clearance and my seven-year-old son declared I needed to hold all of his stuff this week as getting in and out is a ‘two-handed- operation'. Like him, I too am grateful for the grab handles and side steps in this car.

The technology is well-rounded and simple to use. The 12.3-inch touchscreen multimedia system looks great and is responsive to touch. It has built-in satellite navigation and a Toyota Connected Services app with three years of complimentary updates. There is also wired Apple CarPlay and Android Auto, AM/FM radio and DAB+ radio but there’s not much else to access in this system.

The mostly analogue instrument cluster features a 7.0-inch tech screen and this is where you can do a lot of your customisations with display and safety.

The charging options are solid up front with the choice of a USB-A and C port, 12-volt socket and a wireless charging pad. The rear gets two USB-C ports and a 12-volt socket as well.

The boot is cavernous with its 1131L of luggage capacity and that’s with all seats in use. The squared shape of the boot and the 90-degree angle the boot lid opens to are quite practical, especially when fitting bulky items or loading things in the rain.

All GR Sports come with a powered boot lid and a domestic 220-volt socket for charging larger items while adventuring.

Beyond the subtle colour treatment, the interior remains as it is in the HSE variant on which this special edition is based.

In other words, the cabin has a premium look and feel about it, without sacrificing anything in terms of how practical everything is.

It’s a pleasant mix of durable life-friendly materials – carpet mats and soft-touch surfaces – and stylish touches, such as a metal Defender-stamped section in front of the front passenger. 

The leather steering wheel and shifter are standard, but a premium non-leather wheel is a no-cost option. 

Just one example of where JLR gets attention to detail in the Defender so right is the small hard-rubber textured patches on the back of the steering wheel, where your fingertips naturally rest when you’re driving.

It’s those kinds of seemingly minor additions – as well as grippy cargo-area floor surfaces and the like – that add worthwhile tactile elements to the overall Defender package.

Overall, cabin layout is user-friendly with all controls easy to locate and operate.

Storage spaces include a deep centre console, glove box, twin cupholders in between driver and passenger, sunglass storage, door pockets, and shallow spaces peppered around for your pocket gear.

Charge points include USBs up front, and a wireless charging tray.

The driver and front passenger get plenty of room and amenities and those behind them also fare well with adequate room for everyone. 

All seats are very comfortable (can this really be a Defender?) and the front seats are 14-way power-adjustable with heating, cooling and memory.

The second row – a 40/20/40 folding configuration – is heated and has a centre armrest. Second-row passengers have access to air-vent controls and USB charge points in the rear of the centre console.

The rear cargo area seems a bit small in this five-seater, although it offers a listed 1075 litres. With the second row folded down, there is a claimed 2380 litres of space. It has cargo-restraint points.

That area has a sliding cover which conceals your valuables from the prying eyes of nefarious types.

Obviously, there is a lot more to admire here for those who love the interiors of prestige cars, but rather than spending my precious time fondling leather accents or going ‘ohhhh-ahhhh’ over a sunroof or the warming qualities of a heated seat, I like to actually drive. 

And drive I did.

The limited-edition Defenders are based on the high-spec HSE variant and are available in the three-door 90 body style, or the five-door 110 body style. 

Our test vehicle is the 110 and has a recommended retail price of $156,157, excluding on-road costs. 

Each special Defender is a P400 MHEV (mild hybrid electric vehicle), so they have a turbocharged 3.0-litre in-line six-cylinder petrol engine, supported by a small electric motor.

As you’d expect, the standard features list for this 75th Limited Edition is humongous, however, for the sake of brevity, we’ll list only some of the more notable items, which include a 11.4-inch 'Pivi Pro' multi-media system, 3D surround camera, configurable 'Terrain Response' system, Meridian sound system, Matrix LED front lighting, a head-up display, a wireless device charger, as well as 14-way power-adjustable front seats with heating, cooling and memory, a sliding panoramic glass sunroof and three-zone climate control. 

Elements specific to the 75th anniversary edition include an exterior paint finish of 'Grasmere Green', as well as 20-inch alloy wheels in the same colour, with matching centre caps.

There is also a ‘75 years’ graphic, 'Ceres Silver' bumpers and privacy glass. Inside, the Cross Car Beam is finished in brushed Grasmere Green powder coat, the interior is Ebony and there are “Ebony grained leather seat facings”, according to JLR.

Defender buyers may, of course, option up their vehicle with a plethora of accessories and packs, which include grouped accessories to suit your specific lifestyle, i.e. 'Adventure', 'Explorer', 'Country' or 'Urban'.

The LC300 models all share the same 3.3-litre V6 twin-turbo diesel engine that produces a hefty 227kW and 700Nm of torque. Making it powerful enough to easily handle its 3.5-tonne braked towing capacity.

The GR Sport has a 10-speed auto transmission and has a full-time 4WD system with high and low range. It also has front, rear and centre diff locks, so adventuring pursuits should be a breeze but check out Crafty’s off-road review on this model for more 4WD insights.

As mentioned earlier, this Defender is a P400 MHEV (mild hybrid electric vehicle), so it has a 3.0-litre in-line six-cylinder turbo-petrol engine, supported by a small electric motor.

That engine produces 294kW at 5500-6500rpm and 550Nm at 2000-5000rpm and it’s matched to an eight-speed automatic transmission. 

The P400 MHEV has a 48-volt lithium-ion battery, aimed at reducing engine load and fuel consumption, and it has a 7.0kW electric supercharger aimed at minimising turbo lag.

The Defender has permanent all-wheel drive and a dual-range transfer case with high- and low-range 4WD.

It has a comprehensive suite of driver-assist tech, including Land Rover’s 'Terrain Response 2' system, with switchable modes such as 'Grass/Gravel/Snow', 'Sand', 'Mud and Ruts', and 'Rock Crawl'. 

That system optimises throttle response, engine outputs, transmission shifts and diff control to best suit the terrain. It also has centre and rear diff locks.

Our test vehicle also has Land Rover's configurable terrain response system (giving the driver the ability to select and save powertrain, traction and diff settings for off-road driving) and a wade program, which increases air-suspension ride height and closes off the climate control vents to reduce the ingress of moisture in the air.

This Defender has an official 0-100km/h sprint time of 6.0 seconds, and I can confirm that this has plenty of punch off the mark and when overtaking on open roads. It has a listed maximum speed of 191km/h.

It's big and has a lot of power, so it should be pretty thirsty ... right?

Wrong! The official combined fuel-cycle consumption figure is 8.9L/100km and my real world usage came out at 9.8L/100km after doing a mix of long open-road trips and some urban stuff.

Based on the official combined fuel cycle and the two fuel tanks, which equal 110L, you’d be able to get a theoretical driving range of 1236km.

Fuel consumption is a claimed 9.9L/100km (on the combined cycle), and we recorded actual fuel consumption on test of 11L/100km.

The Defender has a 90-litre tank so, going by that fuel-consumption figure, you could reasonably expect a driving range of just under 800km on a full tank, factoring in a safe-distance buffer of 20km; 818km without the buffer.

The GR Sport is a massive car but it doesn’t handle like one. That's not to say it handles like a small car but it doesn't feel like a truck to drive.

The power is effortlessly gutsy and there's no issue keeping your speed consistent on hills. The 10-speed auto transmission manages its gear changes and power hits smoothly. The rumble of the V6 engine is also quite therapeutic when it comes time to tackle an overflowing causeway or muddy road.

The GR Sport feels solidly grounded and even on winding roads, the roll in corners isn’t bad at all. Less so than a Prado, actually. 

The ride comfort is very good in the GR Sport grade as it has special adaptive suspension and that means that you really aren’t bothered by the road surface. Besides some whistling from the roof racks, there’s also little outside noise and it feels refined in the cabin because of it.

The wide windows and high seating position offers great visibility all around but I'm very aware of how large the car is as I can't see my son when he walks around it. The sensors alert you but make sure all kids are accounted for before you start moving.

Despite its size, the GR Sport is very nimble to manoeuvre with a small 11.8m turning circle and steering that is responsive.

Parking has been no issue for me this week and while the 360-degree view camera is a bit too fish-eye lense for me, its clear and you get used to it pretty quickly.

The Defender’s 75th anniversary touches are all cosmetic which is fine because, as is, this Landie is surprisingly impressive on-road, and supremely effective off-road.

This 110 variant is 5018mm long (including the rear-mounted spare tyre), 2008mm wide and 1972mm high with a 3022mm-long wheelbase.

It has a turning circle of 12.8m and a kerb weight is 2297kg. 

So, it’s not a small vehicle, but it never feels unwieldy to drive and it even manages to consistently feel lively and dynamic.

The Defender has a real planted feel on the road; it’s composed and very comfortable, no matter how hard you drive it. 

It’s also quiet. Very quiet, and oh-so-refined, with noise, vibration and harshness levels having been subdued to almost nothing.

The new Defender’s cabin is a pleasantly cocooned space, in which you feel insulated from the world around you. There is a bit of wind rush around the Defender’s wing mirrors, but nothing atrocious. 

Throttle response is crisp, and the 3.0-litre petrol’s 294kW and 550Nm are readily available for a punchy standing-start, or to safely and smoothly overtake another vehicle on the highway – or during low-range 4WDing, but more about that later*.

(* Okay, if you can’t wait, skip ahead to read about the off-road section of this test.)

The eight-speed automatic transmission is very cluey – it’s smarter than you and me – but I found the shifter’s size (stubby) and position (just under the multimedia system’s touchscreen) a bit annoying to work with as I preferred to use Sport/manual mode, sometimes requiring quick shifts up or down.

Road-holding is tremendous, especially considering previous Defenders were about as composed as wonky tractors, and comfort levels are exceptional for something that was once considered a form of punishment to travel in.

Our test vehicle was on 20-inch rims and 255/60 R20 Goodyear Wrangler ‘all-terrain adventure’ tyres and those are perfectly fine for driving on blacktop.

However, they became quickly gummed up with clay and mud during our 4WD testing phase, but more about that soon.

So, how does it perform off-road? Very well, thank you very much. It does what older Defenders can do – and more. And it does it all with supreme levels of comfort and composure – something that can’t be said of older Defenders.

In terms of off-road measures, the Defender has a claimed 228mm-291mm of ground clearance (courtesy of height-adjustable air suspension) and a wading depth of 900mm (again, with the benefit of the air suspension).

If the Defender’s under-carriage hits dirt, the air suspension automatically applies an emergency 75mm of extra height.

This Defender has approach, ramp (breakover) and departure angles of 38 degrees, 28 degrees, and 40 degrees, respectively.

So, it well and truly ticks all of those boxes.

On the dirt-track drive to our 4WD testing and proving ground the Defender demonstrated, as it has before, that it’s able to master and make bearable pretty much any road or track surface, no matter what sort of terrible condition that terrain is in.

This Landie was always stable and planted on the track, with plenty of the credit for that going to a robust monocoque chassis, as well as a multi-link set-up and fully independent air suspension, which as a combination works supremely well to smooth out surface imperfections at all speeds. 

Ride and handling at speed through deep-rutted and potholed sections of dirt road was impressively smooth and composed.

With the air suspension raised to off-road height and tyre pressures dropped to 26 psi, we were ready to put the Defender through its paces.

For those of you who haven’t seen any of my videos or read any of my yarns, our testing site offers more than enough of a challenge to push any standard 4WD to the limits of its reasonable use. I’ve even seen modified vehicles struggle on some of our set-piece challenges.

The terrain here is a mix of steep rocky hill-climbs, slippery descents, mud-holes and water-crossings and, depending on the weather, it can either be seriously challenging or pretty bloody dangerous.

No need for any concern though because the Defender did everything asked of it – and did it all with relaxed ease. I’ve driven an all-conquering 90 at this test site before and the 110 this time was just as impressive.

Steering has a light but precise feel about it at low speeds and that's crucial for such a big vehicle during low-range 4WDing, especially when picking your right line on challenges or manoeuvring along tight, twisted tracks.

There’s plenty of low-down torque from the petrol engine and in low-range that’s evenly applied.

The Pivi touchscreen system is the new Defender’s command centre and, using it, you’re able to cycle through the terrain response programs, you can set driving modes, and essentially operate everything. 

It's generally easy enough to use, but it is a bit tricky to operate on the move – and that’s one of the few niggles I have about this Defender.

The terrain response system, which enables the driver to dial through driving modes, including Mud and Ruts, and Rock Crawl is a clever set-up and would certainly feel out of place in an old-school Defender.

You can calibrate the system’s responses – acceleration, traction sensitivity, and diff control – to suit your driving style and the terrain you’re traversing.

The tread of the Defender’s Goodyear Wrangler all-terrain tyres, as mentioned earlier, swiftly became gummed up with sloppy mud and we lost a fair bit in terms of reliable traction from that rubber on the rock surfaces we moved to straight after driving through a series of shallow mud-holes.

Wheel travel is decent with the Defender able to get useable flex out of that multi-link set-up and air suspension combo.

Another of my very few gripes about the Defender is the fact that while all of the off-road-focussed driver-assist tech, especially terrain response, is so seamlessly effective – it’s almost too good for its own good.

As the driver I almost feel removed from the experience of tackling the terrain I’m on.

Driving this doesn't feel like such a visceral experience as it does in the Defenders of old, or even as hands-on as it does when driving off-road in rebooted old-school 4WDs, such as the Suzuki Jimny, or the Jeep Wrangler Rubicon. 

The new Defender is very capable, and comfortably so, but it feels a little bit too clinical and calculated in its execution.

In terms of its potential as a towing platform, the Defender has a claimed maximum unbraked trailer capacity of 750kg and a maximum braked towing capacity of 3500kg.

It has a maximum roof load of 300kg. GVM (gross vehicle mass) is 3165kg and GCM (gross combined mass) is 6665kg.

The Defender range has the maximum five-star ANCAP safety rating from testing conducted in 2020.

It has a stack of safety gear as standard and driver-assist tech includes AEB, adaptive cruise control, driver condition monitor, blind spot assist, lane keep assist, forward traffic detection, a 3D surround camera, 360-degree parking aid, traffic sign recognition and adaptive speed limiter, and more.

It also has tech that comes in useful for on- and off-road tourers including its ‘transparent’ bonnet view, wade sensing, tyre pressure monitoring system, and tow hitch assist.

It has three top tethers for child seats across the second row and ISOFIX points on the outboard positions only.

The GR Sport comes with a five-year/unlimited kilometre warranty and if you stick to your service schedule, you get an additional two years of engine coverage.

There is capped-priced servicing for up to five years or 100,000km, which entails a total of 10 services or two a year as servicing intervals are a pain at every six months or 10,000km, whichever occurs first.

Services are $400 which equates to $800 per year, which is a bit expensive but not outrageous for the class.

The Defender has a five year/unlimited km warranty with five years roadside assistance. Pretty much the expected norm in this part of the market.

Service intervals are set for every five years or 102,000km (maximum), whichever occurs first – and that’s for a total cost of $2250, or an annual average of $450, which is pretty handy.